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Biology · Introductory biology · Worked example

Estimate population size with mark–recapture

Biologists mark 60 turtles in a pond and release them. Two weeks later they catch 75 turtles, 15 of them marked. Estimate the population with the Lincoln–Petersen and Chapman formulas, and say how precise the estimate is.

Set up the proportion

Marked turtles are 15 of the 75 caught, one fifth. If the same fraction of the whole population is marked, the 60 marked turtles are one fifth of N.

1575=60NN=300

Apply the Lincoln–Petersen formula

The same answer comes straight from N̂ = MC/R.

60⋅7515=300

Apply Chapman’s correction

Chapman’s formula gives a slightly smaller, less biased estimate.

61⋅7616−1=288.75

Gauge the uncertainty

For Chapman’s estimate the standard error is about 53.6 turtles, so the approximate 95% interval is 288.75 ± 1.96 × 53.6, about 184 to 394 turtles. Fifteen recaptures pin the population down only roughly.

1.96⁢(53.63)≈105.11

Check the assumptions

The estimate assumes that no turtles were born, died or moved in or out during the two weeks, that marks stayed visible, that marked turtles mixed back in, and that every turtle was equally likely to be caught.

Result

About 300 turtles by Lincoln–Petersen, or 289 by Chapman’s correction; the approximate 95% interval runs from about 184 to 394.

Your turn

A biologist marks 120 fish; a later catch of 100 fish includes 8 marked ones. Estimate the population with the Lincoln–Petersen formula.

Show the answer and explanation

About 1,500 fish; Chapman’s correction gives about 1,357.

N̂ = MC/R = 120 × 100/8 = 1,500. With only 8 recaptures the estimate is very uncertain.

120⋅1008=1500

Keep exploring

The mark–recapture tool opens with these counts and shows the Chapman estimate, its standard error and the approximate interval. It lists each sampling assumption until you confirm it.

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